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Metabolic Pathways in Higher Plants and Their Determination

Metabolites and Their Types

What Are Metabolites?

  • Metabolites are small molecules involved in metabolism, essential for energy production, growth, repair, and various cellular functions.

  • They are intermediates or end products of metabolic pathways.

Types of Metabolites

Primary Metabolites

  • Involved in basic processes like growth and energy production.

  • Examples: Glucose, amino acids, ATP, fatty acids.

Secondary Metabolites

  • Non-essential for growth but important for defense, signaling, and adaptation.

  • Examples: Alkaloids (nicotine), terpenoids (menthol), flavonoids, antibiotics (erythromycin).

Primary Metabolism

  • Primary metabolism involves pathways essential for life, such as photosynthesis, respiration, and the biosynthesis of carbohydrates, amino acids, and lipids.

Key Processes:

Photosynthesis:

  • Converts light energy into chemical energy, forming sugars from carbon dioxide and water.

Respiration:

  • Breaks down sugars to release energy for cellular processes.

Biosynthetic Routes:

  • Carbohydrate Synthesis: Produces sugars via the Calvin Cycle.

  • Amino Acid Metabolism: Forms the building blocks for proteins.

  • Lipid Synthesis: Builds cell membranes and energy-storing molecules.

Secondary Metabolism

  • Secondary metabolites are derived from primary metabolic pathways and include diverse classes of compounds such as alkaloids, terpenoids, and phenolics.

Biosynthetic Origins:

  • Phenylpropanoids: Derived from the amino acid phenylalanine.

  • Terpenoids: Derived from isoprene units originating in carbohydrate metabolism.

  • Alkaloids: Derived from amino acids and other primary metabolic intermediates.

Determination of Metabolic Pathways

  • Understanding metabolic pathways requires experimental techniques that map out the biosynthesis and function of metabolites:

Key Techniques:

Biochemical Techniques:

  • Enzyme isolation, in vitro assays, and substrate feeding experiments to identify enzymatic steps in pathways.

Radioisotope Labeling:

  • Introducing radioactive isotopes (e.g., 14C or 3H) into precursors to track the movement of atoms through metabolic routes.

Analytical Methods:

  • Techniques like chromatography, mass spectrometry, and NMR are used to identify and quantify metabolites.

Molecular Biology Tools:

  • Genetic manipulation, gene expression profiling, and mutant analysis to study enzyme roles and regulatory networks.


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